Deer overpopulation in many parts of Scotland is harming the environment. Attempts to tackle the longstanding issue are costing taxpayers tens of millions of pounds.

Scotland’s publicly-funded forestry bodies have spent more than £134m controlling deer over the last decade, The Ferret can reveal.

The country’s’s deer population is estimated to have reached more than one million, having doubled since the 1990s. In high concentrations, deer can cause numerous issues.

They include the deaths of an estimated 700 people in vehicle collisions in the UK every year, the environmental harm caused by trampling and overgrazing, poor animal welfare due to insufficient food and shelter for deer in winter, and steep economic costs.